President Emmanuel Macron sought to calm flaring tensions with Muslims around the world on Saturday, telling an Arab TV channel he understood that caricatures of the Prophet Mohammed could be shocking while lashing out at "lies" that the French state was behind them.

France is on edge after the republication in early September of cartoons of the prophet by the Charlie Hebdo weekly, which was followed by an attack outside its former offices, the beheading of a teacher and an attack on a church in Nice Thursday that left three dead.

The country was further shaken by a new incident on Saturday, when an attacker armed with a sawn-off shotgun shot an Orthodox priest as he closed his church in the French city of Lyon before fleeing, a police source said.

Police block access to the Notre-Dame de l'Assomption Basilica in Nice where a knife-wielding young Tunisian killed three people in an attack the authorities are treating as an act of "Islamist terror"
Police block access to the Notre-Dame de l'Assomption Basilica in Nice where a knife-wielding young Tunisian killed three people in an attack the authorities are treating as an act of "Islamist terror" AFP / Valery HACHE

The 52-year-old priest, who has Greek nationality, was shot in the liver at point-blank range and taken to hospital in a serious condition, sources said.

A suspect was arrested later Saturday, Lyon's public prosecutor said, with the motive of the attack remaining unclear.

Muslim devotees burn a French national flag during a demonstration to protest against French President Emmanuel Macron in India.
Muslim devotees burn a French national flag during a demonstration to protest against French President Emmanuel Macron in India. AFP / NARINDER NANU

Macron sparked protests across the Muslim world after the murder earlier this month of teacher Samuel Paty -- who had shown his class a cartoon of Mohammed -- by saying France would never renounce its laws permitting blasphemous caricatures.

But in an apparent bid to reach out to Muslims, Macron gave a long interview setting out his vision to Qatar-based TV channel Al-Jazeera, seeking to strike a softer tone.

Macron said that 'I can understand that people could be shocked by the caricatures.'
Macron said that 'I can understand that people could be shocked by the caricatures.' POOL / Lewis Joly

"I can understand that people could be shocked by the caricatures, but I will never accept that violence can be justified," he said.

"I understand the feelings that this arouses, I respect them. But I want you to understand the role that I have. My role is to calm things down, as I am doing here, but at the same time it is to protect these rights."

He added: "I will always defend in my country the freedom to speak, to write, to think, to draw."

Around 10,000 people marched through Karachi, Pakistan's biggest city, after Friday prayers in what was organised as a procession to mark the Prophet's birthday, but which was charged with anti-France anger. French President Emmanuel Macron has vowed to
Around 10,000 people marched through Karachi, Pakistan's biggest city, after Friday prayers in what was organised as a procession to mark the Prophet's birthday, but which was charged with anti-France anger. French President Emmanuel Macron has vowed to take the fight to religious radicals after the October 16 beheading of a history teacher by an extremist for having shown pupils cartoons of the Prophet Mohammed during a lesson on free speech. AFPTV / Israr AHMED KHAN

Macron lashed out at "distortions" from political leaders over the cartoons of the prophet, saying too often people were led to believe that they were a creation of the French state.

Around 10,000 people marched through Karachi, Pakistan's biggest city, after Friday prayers in what was organised as a procession to mark the Prophet's birthday, but which was charged with anti-France anger. French President Emmanuel Macron has vowed to
Around 10,000 people marched through Karachi, Pakistan's biggest city, after Friday prayers in what was organised as a procession to mark the Prophet's birthday, but which was charged with anti-France anger. French President Emmanuel Macron has vowed to take the fight to religious radicals after the October 16 beheading of a history teacher by an extremist for having shown pupils cartoons of the Prophet Mohammed during a lesson on free speech. AFPTV / Israr AHMED KHAN

He slammed "a confusion that has been fed by many media -- and sometimes political and religious leaders -- which is to say that these caricatures are in a way the project or the creation of the French government or the president".

He also denounced calls for a boycott of French goods, backed in particular by Turkish President Recep Tayyip Erdogan and taken up by some retailers in Muslim countries, as "unworthy" and "unacceptable".

He said the campaign was created by some private groups "who relied on lies... sometimes from other leaders" about the caricatures.

Even before the attack on Paty, Macron had promised a tough new campaign against Islamist radicalism in France, which has claimed the lives of hundreds of people since 2015.

Protests erupted Friday in Afghanistan, Pakistan, Bangladesh, Mali, Mauritania and Lebanon, the latest in a string of mass rallies denouncing France.

France is still reeling from the latest attack in Nice, which Macron has already described as "Islamist" terror.

French authorities were on Saturday seeking to ascertain if a young Tunisian suspected of killing three people in a knife rampage inside a Nice church had outside help.

Brahim Issaoui, 21, only arrived in Europe from Tunisia last month and, according to prosecutors, killed a church employee, a Brazilian woman and a French woman in the attack in the Notre-Dame Basilica on Thursday morning.

The attacker cut the throat of Nadine Devillers, 60, and the sexton Vincent Loques, 55. A Brazilian mother, Simone Barreto Silva, who was stabbed several times, took refuge in a nearby restaurant but died of her wounds there.

Issaoui was shot by police multiple times and is currently in a grave condition in hospital. Investigators have been unable to question him and his precise motivations remain unclear.

"It is still too early to say if there were others complicit, what his motivations were in coming to France and when this idea took root in him," said a source close to the inquiry who asked not to be named.

Investigators believe Issaoui travelled illegally to Europe via Italy's Mediterranean island of Lampedusa on September 20.

He arrived at the mainland Italian port of Bari on October 9 before coming to Nice just one or two days before the attack.